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We propose to determine if the control of copepod populations in the spring and fall are due to predation by gelatinous zooplankton (top-down control), or due to food limitations (bottom-up control). We will measure predation rates on copepods and ctenophores by gelatinous species, biomass of gelatinous and crustacean zooplankton, copepod grazing rates on phytoplankton, protozoa and detritus, and copepod reproduction.